Chapter III Struggle for
Existence
Before entering on the subject of this chapter, I
must make a few preliminary remarks, to show how the struggle for existence
bears on Natural Selection. It has been seen in the last chapter that amongst
organic beings in a state of nature there is some individual variability;
indeed I am not aware that this has ever been disputed. It is immaterial for
us whether a multitude of doubtful forms be called species or sub-species or
varieties; what rank, for instance, the two or three hundred doubtful forms of
British plants are entitled to hold, if the existence of any well-marked
varieties be admitted. But the mere existence of individual variability and of
some few well-marked varieties, though necessary as the foundation for the
work, helps us but little in understanding how species arise in nature. How
have all those exquisite adaptations of one part of the organisation to
another part, and to the conditions of life, and of one distinct organic being
to another being, been perfected? We see these beautiful co-adaptations most
plainly in the woodpecker and missletoe; and only a little less plainly in the
humblest parasite which clings to the hairs of a quadruped or feathers of a
bird; in the structure of the beetle which dives through the water; in the
plumed seed which is wafted by the gentlest breeze; in short, we see beautiful
adaptations everywhere and in every part of the organic world.
2 Again, it may be asked, how is it that varieties,
which I have called incipient species, become ultimately converted into good
and distinct species, which in most cases obviously differ from each other far
more than do the varieties of the same species? How do those groups of
species, which constitute what are called distinct genera, and which differ
from each other more than do the species of the same genus, arise? All these
results, as we shall more fully see in the next chapter, follow inevitably
from the struggle for life. Owing to this struggle for life, any variation,
however slight and from whatever cause proceeding, if it be in any degree
profitable to an individual of any species, in its infinitely complex
relations to other organic beings and to external nature, will tend to the
preservation of that individual, and will generally be inherited by its
offspring. The offspring, also, will thus have a better chance of surviving,
for, of the many individuals of any species which are periodically born, but a
small number can survive. I have called this principle, by which each slight
variation, if useful, is preserved, by the term of Natural Selection, in order
to mark its relation to man's power of selection. We have seen that man by
selection can certainly produce great results, and can adapt organic beings to
his own uses, through the accumulation of slight but useful variations, given
to him by the hand of Nature. But Natural Selection, as we shall hereafter
see, is a power incessantly ready for action, and is as immeasurably superior
to man's feeble efforts, as the works of Nature are to those of Art.
3 We will now discuss in a little more detail the
struggle for existence. In my future work this subject shall be treated, as it
well deserves, at much greater length. The elder De Candolle and Lyell have
largely and philosophically shown that all organic beings are exposed to
severe competition. In regard to plants, no one has treated this subject with
more spirit and ability than W. Herbert, Dean of Manchester, evidently the
result of his great horticultural knowledge. Nothing is easier than to admit
in words the truth of the universal struggle for life, or more difficult--at
least I have found it so--than constantly to bear this conclusion in mind. Yet
unless it be thoroughly engrained in the mind, I am convinced that the whole
economy of nature, with every fact on distribution, rarity, abundance,
extinction, and variation, will be dimly seen or quite misunderstood. We
behold the face of nature bright with gladness, we often see superabundance of
food; we do not see, or we forget, that the birds which are idly singing round
us mostly live on insects or seeds, and are thus constantly destroying life;
or we forget how largely these songsters, or their eggs, or their nestlings,
are destroyed by birds and beasts of prey; we do not always bear in mind, that
though food may be now superabundant, it is not so at all seasons of each
recurring year.
4 I should premise that I use the term Struggle for
Existence in a large and metaphorical sense, including dependence of one being
on another, and including (which is more important) not only the life of the
individual, but success in leaving progeny. Two canine animals in a time of
dearth, may be truly said to struggle with each other which shall get food and
live. But a plant on the edge of a desert is said to struggle for life against
the drought, though more properly it should be said to be dependent on the
moisture. A plant which annually produces a thousand seeds, of which on an
average only one comes to maturity, may be more truly said to struggle with
the plants of the same and other kinds which already clothe the ground. The
missletoe is dependent on the apple and a few other trees, but can only in a
far-fetched sense be said to struggle with these trees, for if too many of
these parasites grow on the same tree, it will languish and die. But several
seedling missletoes, growing close together on the same branch, may more truly
be said to struggle with each other. As the missletoe is disseminated by
birds, its existence depends on birds; and it may metaphorically be said to
struggle with other fruit-bearing plants, in order to tempt birds to devour
and thus disseminate its seeds rather than those of other plants. In these
several senses, which pass into each other, I use for convenience sake the
general term of struggle for existence.
5 A struggle for existence inevitably follows from the
high rate at which all organic beings tend to increase. Every being, which
during its natural lifetime produces several eggs or seeds, must suffer
destruction during some period of its life, and during some season or
occasional year, otherwise, on the principle of geometrical increase, its
numbers would quickly become so inordinately great that no country could
support the product. Hence, as more individuals are produced than can possibly
survive, there must in every case be a struggle for existence, either one
individual with another of the same species, or with the individuals of
distinct species, or with the physical conditions of life. It is the doctrine
of Malthus applied with manifold force to the whole animal and vegetable
kingdoms; for in this case there can be no artificial increase of food, and no
prudential restraint from marriage. Although some species may be now
increasing, more or less rapidly, in numbers, all cannot do so, for the world
would not hold them.
6 There is no exception to the rule that every organic
being naturally increases at so high a rate, that if not destroyed, the earth
would soon be covered by the progeny of a single pair. Even slow-breeding man
has doubled in twenty-five years, and at this rate, in a few thousand years,
there would literally not be standing room for his progeny. Linnaeus has
calculated that if an annual plant produced only two seeds--and there is no
plant so unproductive as this--and their seedlings next year produced two, and
so on, then in twenty years there would be a million plants. The elephant is
reckoned to be the slowest breeder of all known animals, and I have taken some
pains to estimate its probable minimum rate of natural increase: it will be
under the mark to assume that it breeds when thirty years old, and goes on
breeding till ninety years old, bringing forth three pair of young in this
interval; if this be so, at the end of the fifth century there would be alive
fifteen million elephants, descended from the first pair.
7 But we have better evidence on this subject than mere
theoretical calculations, namely, the numerous recorded cases of the
astonishingly rapid increase of various animals in a state of nature, when
circumstances have been favourable to them during two or three following
seasons. Still more striking is the evidence from our domestic animals of many
kinds which have run wild in several parts of the world: if the statements of
the rate of increase of slow-breeding cattle and horses in South America, and
latterly in Australia, had not been well authenticated, they would have been
quite incredible. So it is with plants: cases could be given of introduced
plants which have become common throughout whole islands in a period of less
than ten years. Several of the plants now most numerous over the wide plains
of La Plata, clothing square leagues of surface almost to the exclusion of all
other plants, have been introduced from Europe; and there are plants which now
range in India, as I hear from Dr. Falconer, from Cape Comorin to the Himalaya,
which have been imported from America since its discovery. In such cases, and
endless instances could be given, no one supposes that the fertility of these
animals or plants has been suddenly and temporarily increased in any sensible
degree. The obvious explanation is that the conditions of life have been very
favourable, and that there has consequently been less destruction of the old
and young, and that nearly all the young have been enabled to breed. In such
cases the geometrical ratio of increase, the result of which never fails to be
surprising, simply explains the extraordinarily rapid increase and wide
diffusion of naturalised productions in their new homes.
8 In a state of nature almost every plant produces
seed, and amongst animals there are very few which do not annually pair. Hence
we may confidently assert, that all plants and animals are tending to increase
at a geometrical ratio, that all would most rapidly stock every station in
which they could any how exist, and that the geometrical tendency to increase
must be checked by destruction at some period of life. Our familiarity with
the larger domestic animals tends, I think, to mislead us: we see no great
destruction falling on them, and we forget that thousands are annually
slaughtered for food, and that in a state of nature an equal number would have
somehow to be disposed of.
9 The only difference between organisms which annually
produce eggs or seeds by the thousand, and those which produce extremely few,
is, that the slow-breeders would require a few more years to people, under
favourable conditions, a whole district, let it be ever so large. The condor
lays a couple of eggs and the ostrich a score, and yet in the same country the
condor may be the more numerous of the two: the Fulmar petrel lays but one
egg, yet it is believed to be the most numerous bird in the world. One fly
deposits hundreds of eggs, and another, like the hippobosca, a single one; but
this difference does not determine how many individuals of the two species can
be supported in a district. A large number of eggs is of some importance to
those species, which depend on a rapidly fluctuating amount of food, for it
allows them rapidly to increase in number. But the real importance of a large
number of eggs or seeds is to make up for much destruction at some period of
life; and this period in the great majority of cases is an early one. If an
animal can in any way protect its own eggs or young, a small number may be
produced, and yet the average stock be fully kept up; but if many eggs or
young are destroyed, many must be produced, or the species will become
extinct. It would suffice to keep up the full number of a tree, which lived on
an average for a thousand years, if a single seed were produced once in a
thousand years, supposing that this seed were never destroyed, and could be
ensured to germinate in a fitting place. So that in all cases, the average
number of any animal or plant depends only indirectly on the number of its
eggs or seeds.
10 In looking at Nature, it is most necessary to keep
the foregoing considerations always in mind--never to forget that every single
organic being around us may be said to be striving to the utmost to increase
in numbers; that each lives by a struggle at some period of its life; that
heavy destruction inevitably falls either on the young or old, during each
generation or at recurrent intervals. Lighten any check, mitigate the
destruction ever so little, and the number of the species will almost
instantaneously increase to any amount. The face of Nature may be compared to
a yielding surface, with ten thousand sharp wedges packed close together and
driven inwards by incessant blows, sometimes one wedge being struck, and then
another with greater force.
11 What checks the natural tendency of each species to
increase in number is most obscure. Look at the most vigorous species; by as
much as it swarms in numbers, by so much will its tendency to increase be
still further increased. We know not exactly what the checks are in even one
single instance. Nor will this surprise any one who reflects how ignorant we
are on this head, even in regard to mankind, so incomparably better known than
any other animal. This subject has been ably treated by several authors, and I
shall, in my future work, discuss some of the checks at considerable length,
more especially in regard to the feral animals of South America. Here I will
make only a few remarks, just to recall to the reader's mind some of the chief
points. Eggs or very young animals seem generally to suffer most, but this is
not invariably the case. With plants there is a vast destruction of seeds,
but, from some observations which I have made, I believe that it is the
seedlings which suffer most from germinating in ground already thickly stocked
with other plants. Seedlings, also, are destroyed in vast numbers by various
enemies; for instance, on a piece of ground three feet long and two wide, dug
and cleared, and where there could be no choking from other plants, I marked
all the seedlings of our native weeds as they came up, and out of the 357 no
less than 295 were destroyed, chiefly by slugs and insects. If turf which has
long been mown, and the case would be the same with turf closely browsed by
quadrupeds, be let to grow, the more vigorous plants gradually kill the less
vigorous, though fully grown, plants: thus out of twenty species growing on a
little plot of turf (three feet by four) nine species perished from the other
species being allowed to grow up freely.
12 The amount of food for each species of course gives
the extreme limit to which each can increase; but very frequently it is not
the obtaining food, but the serving as prey to other animals, which determines
the average numbers of a species. Thus, there seems to be little doubt that
the stock of partridges, grouse, and hares on any large estate depends chiefly
on the destruction of vermin. If not one head of game were shot during the
next twenty years in England, and, at the same time, if no vermin were
destroyed, there would, in all probability, be less game than at present,
although hundreds of thousands of game animals are now annually killed. On the
other hand, in some cases, as with the elephant and rhinoceros, none are
destroyed by beasts of prey: even the tiger in India most rarely dares to
attack a young elephant protected by its dam.
13 Climate plays an important part in determining the
average numbers of a species, and periodical seasons of extreme cold or
drought, I believe to be the most effective of all checks. I estimated that
the winter of 1854-55 destroyed four-fifths of the birds in my own grounds;
and this is a tremendous destruction, when we remember that ten per cent. is
an extraordinarily severe mortality from epidemics with man. The action of
climate seems at first sight to be quite independent of the struggle for
existence; but in so far as climate chiefly acts in reducing food, it brings
on the most severe struggle between the individuals, whether of the same or of
distinct species, which subsist on the same kind of food. Even when climate,
for instance extreme cold, acts directly, it will be the least vigorous, or
those which have got least food through the advancing winter, which will
suffer most. When we travel from south to north, or from a damp region to a
dry, we invariably see some species gradually getting rarer and rarer, and
finally disappearing; and the change of climate being conspicuous, we are
tempted to attribute the whole effect to its direct action. But this is a very
false view: we forget that each species, even where it most abounds, is
constantly suffering enormous destruction at some period of its life, from
enemies or from competitors for the same place and food; and if these enemies
or competitors be in the least degree favoured by any slight change of
climate, they will increase in numbers, and, as each area is already fully
stocked with inhabitants, the other species will decrease. When we travel
southward and see a species decreasing in numbers, we may feel sure that the
cause lies quite as much in other species being favoured, as in this one being
hurt. So it is when we travel northward, but in a somewhat lesser degree, for
the number of species of all kinds, and therefore of competitors, decreases
northwards; hence in going northward, or in ascending a mountain, we far
oftener meet with stunted forms, due to the directly injurious action of
climate, than we do in proceeding southwards or in descending a mountain. When
we reach the Arctic regions, or snow-capped summits, or absolute deserts, the
struggle for life is almost exclusively with the elements.
14 That climate acts in main part indirectly by
favouring other species, we may clearly see in the prodigious number of plants
in our gardens which can perfectly well endure our climate, but which never
become naturalised, for they cannot compete with our native plants, nor resist
destruction by our native animals.
15 When a species, owing to highly favourable
circumstances, increases inordinately in numbers in a small tract,
epidemics--at least, this seems generally to occur with our game
animals--often ensue: and here we have a limiting check independent of the
struggle for life. But even some of these so-called epidemics appear to be due
to parasitic worms, which have from some cause, possibly in part through
facility of diffusion amongst the crowded animals, been disproportionably
favoured: and here comes in a sort of struggle between the parasite and its
prey.
16 On the other hand, in many cases, a large stock of
individuals of the same species, relatively to the numbers of its enemies, is
absolutely necessary for its preservation. Thus we can easily raise plenty of
corn and rape-seed, &c., in our fields, because the seeds are in great
excess compared with the number of birds which feed on them; nor can the
birds, though having a superabundance of food at this one season, increase in
number proportionally to the supply of seed, as their numbers are checked
during winter: but any one who has tried, knows how troublesome it is to get
seed from a few wheat or other such plants in a garden; I have in this case
lost every single seed. This view of the necessity of a large stock of the
same species for its preservation, explains, I believe, some singular facts in
nature, such as that of very rare plants being sometimes extremely abundant in
the few spots where they do occur; and that of some social plants being
social, that is, abounding in individuals, even on the extreme confines of
their range. For in such cases, we may believe, that a plant could exist only
where the conditions of its life were so favourable that many could exist
together, and thus save each other from utter destruction. I should add that
the good effects of frequent intercrossing, and the ill effects of close
interbreeding, probably come into play in some of these cases; but on this
intricate subject I will not here enlarge.
17 Many cases are on record showing how complex and
unexpected are the checks and relations between organic beings, which have to
struggle together in the same country. I will give only a single instance,
which, though a simple one, has interested me. In Staffordshire, on the estate
of a relation where I had ample means of investigation, there was a large and
extremely barren heath, which had never been touched by the hand of man; but
several hundred acres of exactly the same nature had been enclosed twenty-five
years previously and planted with Scotch fir. The change in the native
vegetation of the planted part of the heath was most remarkable, more than is
generally seen in passing from one quite different soil to another: not only
the proportional numbers of the heath-plants were wholly changed, but twelve
species of plants (not counting grasses and carices) flourished in the
plantations, which could not be found on the heath. The effect on the insects
must have been still greater, for six insectivorous birds were very common in
the plantations, which were not to be seen on the heath; and the heath was
frequented by two or three distinct insectivorous birds. Here we see how
potent has been the effect of the introduction of a single tree, nothing
whatever else having been done, with the exception that the land had been
enclosed, so that cattle could not enter. But how important an element
enclosure is, I plainly saw near Farnham, in Surrey. Here there are extensive
heaths, with a few clumps of old Scotch firs on the distant hill-tops: within
the last ten years large spaces have been enclosed, and self-sown firs are now
springing up in multitudes, so close together that all cannot live.
18 When I ascertained that these young trees had not
been sown or planted, I was so much surprised at their numbers that I went to
several points of view, whence I could examine hundreds of acres of the
unenclosed heath, and literally I could not see a single Scotch fir, except
the old planted clumps. But on looking closely between the stems of the heath,
I found a multitude of seedlings and little trees, which had been perpetually
browsed down by the cattle. In one square yard, at a point some hundreds yards
distant from one of the old clumps, I counted thirty-two little trees; and one
of them, judging from the rings of growth, had during twenty-six years tried
to raise its head above the stems of the heath, and had failed. No wonder
that, as soon as the land was enclosed, it became thickly clothed with
vigorously growing young firs. Yet the heath was so extremely barren and so
extensive that no one would ever have imagined that cattle would have so
closely and effectually searched it for food.
19 Here we see that cattle absolutely determine the
existence of the Scotch fir; but in several parts of the world insects
determine the existence of cattle. Perhaps Paraguay offers the most curious
instance of this; for here neither cattle nor horses nor dogs have ever run
wild, though they swarm southward and northward in a feral state; and Azara
and Rengger have shown that this is caused by the greater number in Paraguay
of a certain fly, which lays its eggs in the navels of these animals when
first born. The increase of these flies, numerous as they are, must be
habitually checked by some means, probably by birds. Hence, if certain
insectivorous birds (whose numbers are probably regulated by hawks or beasts
of prey) were to increase in Paraguay, the flies would decrease--then cattle
and horses would become feral, and this would certainly greatly alter (as
indeed I have observed in parts of South America) the vegetation: this again
would largely affect the insects; and this, as we just have seen in
Staffordshire, the insectivorous birds, and so onwards in ever-increasing
circles of complexity. We began this series by insectivorous birds, and we
have ended with them. Not that in nature the relations can ever be as simple
as this. Battle within battle must ever be recurring with varying success; and
yet in the long-run the forces are so nicely balanced, that the face of nature
remains uniform for long periods of time, though assuredly the merest trifle
would often give the victory to one organic being over another. Nevertheless
so profound is our ignorance, and so high our presumption, that we marvel when
we hear of the extinction of an organic being; and as we do not see the cause,
we invoke cataclysms to desolate the world, or invent laws on the duration of
the forms of life!
20 I am tempted to give one more instance showing how
plants and animals, most remote in the scale of nature, are bound together by
a web of complex relations. I shall hereafter have occasion to show that the
exotic Lobelia fulgens, in this part of England, is never visited by insects,
and consequently, from its peculiar structure, never can set a seed. Many of
our orchidaceous plants absolutely require the visits of moths to remove their
pollen-masses and thus to fertilise them. I have, also, reason to believe that
humble-bees are indispensable to the fertilisation of the heartsease (Viola
tricolor), for other bees do not visit this flower. From experiments which I
have tried, I have found that the visits of bees, if not indispensable, are at
least highly beneficial to the fertilisation of our clovers; but humble-bees
alone visit the common red clover (Trifolium pratense), as other bees cannot
reach the nectar. Hence I have very little doubt, that if the whole genus of
humble-bees became extinct or very rare in England, the heartsease and red
clover would become very rare, or wholly disappear. The number of humble-bees
in any district depends in a great degree on the number of field-mice, which
destroy their combs and nests; and Mr. H. Newman, who has long attended to the
habits of humble-bees, believes that 'more than two thirds of them are thus
destroyed all over England.' Now the number of mice is largely dependent, as
every one knows, on the number of cats; and Mr. Newman says, 'Near villages
and small towns I have found the nests of humble-bees more numerous than
elsewhere, which I attribute to the number of cats that destroy the mice.'
Hence it is quite credible that the presence of a feline animal in large
numbers in a district might determine, through the intervention first of mice
and then of bees, the frequency of certain flowers in that district!
21 In the case of every species, many different checks,
acting at different periods of life, and during different seasons or years,
probably come into play; some one check or some few being generally the most
potent, but all concurring in determining the average number or even the
existence of the species. In some cases it can be shown that widely-different
checks act on the same species in different districts. When we look at the
plants and bushes clothing an entangled bank, we are tempted to attribute
their proportional numbers and kinds to what we call chance. But how false a
view is this! Every one has heard that when an American forest is cut down, a
very different vegetation springs up; but it has been observed that the trees
now growing on the ancient Indian mounds, in the Southern United States,
display the same beautiful diversity and proportion of kinds as in the
surrounding virgin forests. What a struggle between the several kinds of trees
must here have gone on during long centuries, each annually scattering its
seeds by the thousand; what war between insect and insect--between insects,
snails, and other animals with birds and beasts of prey--all striving to
increase, and all feeding on each other or on the trees or their seeds and
seedlings, or on the other plants which first clothed the ground and thus
checked the growth of the trees! Throw up a handful of feathers, and all must
fall to the ground according to definite laws; but how simple is this problem
compared to the action and reaction of the innumerable plants and animals
which have determined, in the course of centuries, the proportional numbers
and kinds of trees now growing on the old Indian ruins!
22 The dependency of one organic being on another, as of
a parasite on its prey, lies generally between beings remote in the scale of
nature. This is often the case with those which may strictly be said to
struggle with each other for existence, as in the case of locusts and
grass-feeding quadrupeds. But the struggle almost invariably will be most
severe between the individuals of the same species, for they frequent the same
districts, require the same food, and are exposed to the same dangers. In the
case of varieties of the same species, the struggle will generally be almost
equally severe, and we sometimes see the contest soon decided: for instance,
if several varieties of wheat be sown together, and the mixed seed be resown,
some of the varieties which best suit the soil or climate, or are naturally
the most fertile, will beat the others and so yield more seed, and will
consequently in a few years quite supplant the other varieties. To keep up a
mixed stock of even such extremely close varieties as the variously coloured
sweet-peas, they must be each year harvested separately, and the seed then
mixed in due proportion, otherwise the weaker kinds will steadily decrease in
numbers and disappear. So again with the varieties of sheep: it has been
asserted that certain mountain-varieties will starve out other
mountain-varieties, so that they cannot be kept together. The same result has
followed from keeping together different varieties of the medicinal leech. It
may even be doubted whether the varieties of any one of our domestic plants or
animals have so exactly the same strength, habits, and constitution, that the
original proportions of a mixed stock could be kept up for half a dozen
generations, if they were allowed to struggle together, like beings in a state
of nature, and if the seed or young were not annually sorted.
23 As species of the same genus have usually, though by
no means invariably, some similarity in habits and constitution, and always in
structure, the struggle will generally be more severe between species of the
same genus, when they come into competition with each other, than between
species of distinct genera. We see this in the recent extension over parts of
the United States of one species of swallow having caused the decrease of
another species. The recent increase of the missel-thrush in parts of Scotland
has caused the decrease of the song-thrush. How frequently we hear of one
species of rat taking the place of another species under the most different
climates! In Russia the small Asiatic cockroach has everywhere driven before
it its great congener. One species of charlock will supplant another, and so
in other cases. We can dimly see why the competition should be most severe
between allied forms, which fill nearly the same place in the economy of
nature; but probably in no one case could we precisely say why one species has
been victorious over another in the great battle of life.
24 A corollary of the highest importance may be deduced
from the foregoing remarks, namely, that the structure of every organic being
is related, in the most essential yet often hidden manner, to that of all
other organic beings, with which it comes into competition for food or
residence, or from which it has to escape, or on which it preys. This is
obvious in the structure of the teeth and talons of the tiger; and in that of
the legs and claws of the parasite which clings to the hair on the tiger's
body. But in the beautifully plumed seed of the dandelion, and in the
flattened and fringed legs of the water-beetle, the relation seems at first
confined to the elements of air and water. Yet the advantage of plumed seeds
no doubt stands in the closest relation to the land being already thickly
clothed by other plants; so that the seeds may be widely distributed and fall
on unoccupied ground. In the water-beetle, the structure of its legs, so well
adapted for diving, allows it to compete with other aquatic insects, to hunt
for its own prey, and to escape serving as prey to other animals.
25 The store of nutriment laid up within the seeds of
many plants seems at first sight to have no sort of relation to other plants.
But from the strong growth of young plants produced from such seeds (as peas
and beans), when sown in the midst of long grass, I suspect that the chief use
of the nutriment in the seed is to favour the growth of the young seedling,
whilst struggling with other plants growing vigorously all around.
26 Look at a plant in the midst of its range, why does
it not double or quadruple its numbers? We know that it can perfectly well
withstand a little more heat or cold, dampness or dryness, for elsewhere it
ranges into slightly hotter or colder, damper or drier districts. In this case
we can clearly see that if we wished in imagination to give the plant the
power of increasing in number, we should have to give it some advantage over
its competitors, or over the animals which preyed on it. On the confines of
its geographical range, a change of constitution with respect to climate would
clearly be an advantage to our plant; but we have reason to believe that only
a few plants or animals range so far, that they are destroyed by the rigour of
the climate alone. Not until we reach the extreme confines of life, in the
arctic regions or on the borders of an utter desert, will competition cease.
The land may be extremely cold or dry, yet there will be competition between
some few species, or between the individuals of the same species, for the
warmest or dampest spots.
27 Hence, also, we can see that when a plant or animal
is placed in a new country amongst new competitors, though the climate may be
exactly the same as in its former home, yet the conditions of its life will
generally be changed in an essential manner. If we wished to increase its
average numbers in its new home, we should have to modify it in a different
way to what we should have done in its native country; for we should have to
give it some advantage over a different set of competitors or enemies.
28 It is good thus to try in our imagination to give any
form some advantage over another. Probably in no single instance should we
know what to do, so as to succeed. It will convince us of our ignorance on the
mutual relations of all organic beings; a conviction as necessary, as it seems
to be difficult to acquire. All that we can do, is to keep steadily in mind
that each organic being is striving to increase at a geometrical ratio; that
each at some period of its life, during some season of the year, during each
generation or at intervals, has to struggle for life, and to suffer great
destruction. When we reflect on this struggle, we may console ourselves with
the full belief, that the war of nature is not incessant, that no fear is
felt, that death is generally prompt, and that the vigorous, the healthy, and
the happy survive and multiply.