THE ORIGIN OF SPECIES
Charles Darwin

TABLE OF CONTENTS

- Introduction

Chapter I Variation under Domestication

- Causes of Variability
- Effects of Habit
- Correlation of Growth
- Inheritance
- Character of Domestic Varieties
- Difficulty of distinguishing between Varieties and Species
- Origin of Domestic Varieties from one or more Species
- Domestic Pigeons, their Differences and Origin
- Principle of Selection anciently followed, its Effects
- Methodical and Unconscious Selection
- Unknown Origin of our Domestic Productions
- Circumstances favourable to Man's power of Selection.

Chapter II Variation under Nature

- Variability
- Individual Differences
- Doubtful species
- Wide ranging, much diffused, and common species vary most
- Species of the larger genera in any country vary more than the species of the smaller genera
- Many of the species of the larger genera resemble varieties in being very closely, but unequally, related to each other, and in having restricted ranges.

Chapter III Struggle for Existence

- Bears on natural selection
- The term used in a wide sense
- Geometrical powers of increase
- Rapid increase of naturalised animals and plants
- Nature of the checks to increase
- Competition universal
- Effects of climate
- Protection from the number of individuals
- Complex relations of all animals and plants throughout nature
- Struggle for life most severe between individuals and varieties of the same species; often severe between species of the same genus
- The relation of organism to organism the most important of all relations.

Chapter IV Natural Selection

Natural Selection:
        its power compared with man's selection
        its power on characters of trifling importance
        its power at all ages and on both sexes
- Sexual Selection
- On the generality of intercrosses between individuals of the same species --
- Circumstances favourable and unfavourable to Natural Selection, namely, intercrossing, isolation, number of individuals
- Slow action
- Extinction caused by Natural Selection
- Divergence of Character, related to the diversity of inhabitants of any small area, and to naturalisation
- Action of Natural Selection, through Divergence of Character and Extinction, on the    descendants from a common parent
- Explains the Grouping of all organic beings.

Chapter V Laws of Variation

- Effects of external conditions
- Use and disuse, combined with natural selection; organs of flight and of vision --
- Acclimatisation
- Correlation of growth
- Compensation and economy of growth
- False correlations
- Multiple, rudimentary, and lowly organised structures variable
- Parts developed in an unusual manner are highly variable: specific characters more variable than generic:
        secondary sexual characters variable
- Species of the same genus vary in an analogous manner
- Reversions to long-lost characters
- Summary.

Chapter VI Difficulties on Theory

- Difficulties on the theory of descent with modification
- Transitions
- Absence or rarity of transitional varieties
- Transitions in habits of life
- Diversified habits in the same species
- Species with habits widely different from those of their allies
- Organs of extreme perfection
- Means of transition
- Cases of difficulty
- Natura non facit saltum
- Organs of small importance
- Organs not in all cases absolutely perfect
- The law of Unity of Type and of the Conditions of Existence embraced by the theory of Natural Selection.

Chapter VII Instinct

- Instincts comparable with habits, but different in their origin
- Instincts graduated
- Aphides and ants
- Instincts variable
- Domestic instincts, their origin 
- Natural instincts of the cuckoo, ostrich, and parasitic bees
- Slave-making ants
- Hive-bee, its cell-making instinct 
- Difficulties on the theory of the Natural Selection of instincts
- Neuter or sterile insects
- Summary

Chapter VIII Hybridism

- Distinction between the sterility of first crosses and of hybrids
- Sterility various in degree, not universal, affected by close interbreeding, removed by domestication
- Laws governing the sterility of hybrids
- Sterility not a special endowment, but incidental on other differences
- Causes of the sterility of first crosses and of hybrids
- Parallelism between the effects of changed conditions of life and crossing
- Fertility of varieties when crossed and of their mongrel offspring not universal
- Hybrids and mongrels compared independently of their fertility
- Summary

Chapter IX On the Imperfection of the Geological Record

- On the absence of intermediate varieties at the present day
- On the nature of extinct intermediate varieties; on their number
- On the vast lapse of time, as inferred from the rate of deposition and of denudation
- On the poorness of our palaeontological collections
- On the intermittence of geological formations
- On the absence of intermediate varieties in any one formation
- On the sudden appearance of groups of species
- On their sudden appearance in the lowest known fossiliferous strata

Chapter X On the Geological Succession of Organic Beings

- On the slow and successive appearance of new species
- On their different rates of change
- Species once lost do not reappear
- Groups of species follow the same general rules in their appearance and disappearance as do single species
- On Extinction
- On simultaneous changes in the forms of life throughout the world
- On the affinities of extinct species to each other and to living species
- On the state of development of ancient forms
- On the succession of the same types within the same areas
- Summary of preceding and present chapters

Chapter XI Geographical Distribution

- Present distribution cannot be accounted for by differences in physical conditions
- Importance of barriers
- Affinity of the productions of the same continent
- Centres of creation
- Means of dispersal, by changes of climate and of the level of the land, and by occasional means
- Dispersal during the Glacial period co-extensive with the world.

Chapter XII Geographical Distribution (Continued)

- Distribution of fresh-water productions
- On the inhabitants of oceanic islands
- Absence of Batrachians and of terrestrial Mammals
- On the relation of the inhabitants of islands to those of the nearest mainland
- On colonisation from the nearest source with subsequent modification
- Summary of the last and present chapters

Chapter XIII Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Organs

- Classification, groups subordinate to groups
- Natural system
- Rules and difficulties in classification, explained on the theory of descent with modification --
- Classification of varieties
- Descent always used in classification
- Analogical or adaptive characters
- Affinities, general, complex and radiating
- Extinction separates and defines groups
- Morphology, between members of the same class, between parts of the same individual
- Embryology, laws of, explained by variations not supervening at an early age, and being inherited at a corresponding age
- Rudimentary Organs; their origin explained
- Summary

Chapter XIV Recapitulation and Conclusion

- Recapitulation of the difficulties on the theory of Natural Selection
- Recapitulation of the general and special circumstances in its favor
- Causes of the general belief in the immutability of species
- How far the theory of natural selection may be extended
- Effects of its adoption on the study of Natural history
- Concluding remarks